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The skull grew larger, especially the cranium, while the neck shortened to provide better support for the skull. The increase in size led to the development and elongation of the mobile trunk to provide reach. The number of premolars, incisors and canines decreased.[6] The cheek teeth molars and premolars became larger and more specialized, especially after elephants started to switch from C3-plants to C4-grasses, Elephant nice butt poster which caused their teeth to undergo a three-fold increase in teeth height as well as substantial multiplication of lamellae after about five million years ago. Only in the last million years or so did they return to a diet mainly consisting of C3 trees and shrubs.
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The upper second incisors grew into tusks, which varied in shape from straight, to curved either upward or downward to spiralled, depending on the species. Some proboscideans developed tusks from their lower incisors. Elephants retain certain features from their aquatic ancestry, such as their middle ear anatomy. Several species of proboscideans lived on islands and experienced insular dwarfism. This occurred primarily during the Pleistocene when some elephant populations became isolated by fluctuating sea levels, although dwarf elephants did exist earlier in the Pliocene.
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